Family Pycnonotidae, Bulbuls
The Bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical Asia to Indonesia, and north as far as Japan. A few insular species occur on the tropical islands of the Indian Ocean. There are 160 species in 32 genera. While different species are found in a wide range of habitats, the African species are predominantly found in rainforest, whereas Asian bulbuls are predominantly found in more open areas.
The family Pycnonotidae was introduced by the English zoologist George Robert Gray in 1840 as a subfamily Pycnonotinae of the thrush family Turdidae.
The Arabic word bulbul (بلبل) is sometimes used to refer to the "nightingale" as well as the bulbul, but the English word bulbul refers to the birds discussed in this article.
A few species that were previously considered to be members of the Pycnonotidae have been moved to other families. Several Malagasy species that were formerly placed in the genus Phyllastrephus are now placed in the family Bernieridae.[4][5] In addition, the genus Nicator containing three African species is now placed in a separate family Nicatoridae.
A study published in 2007 by Ulf Johansson and colleagues using three nuclear markers found that the genus Andropadus was non-monophyletic. In the subsequent revision, species were moved to three resurrected genera: Arizelocichla, Stelgidillas and Eurillas. Only the sombre greenbul (Andropadus importunus), was retained in Andropadus. A study by Subir Shakya and Frederick Shelden published in 2017 found that species in the large genus Pycnonotus formed several deeply divergent clades. The genus was split and six genera were resurrected to accommodate these clades.
The family forms two main clades. One clade contains species that are only found in Africa; many of these have greenbul in the common name. The second clade contains mostly Asian species but includes a few species that are found in Africa.
The galleries below have images of the various members of Family Pycnonotidae that I have seen over the years. Simply follow the links on the images below:
Buff-vented Bulbul (Iole crypta) South Thailand
Baker’s Bulbul (Iole cinnamomeoventris) Kaeng Krachan
Finsch’s Bulbul (Iole finschii) Hala Bala Thailand
Grey-eyed Bulbul (Iole propinqua) Thailand
Yellow-browed Bulbul (Acritillas indica) Sri Lanka
Mountain Bulbul (Ixos mcclellandii) Thailand
Streaked Bulbul (Ixos malaccensis) Northern Thailand
Brown-eared Bulbul (Hypsipetes amaurotis) Japan
Square-tailed Black Bulbul (Hypsipetes ganeesa) Sri Lanka
Black Bulbul (Hypsipetes leucocephalus) China, Thailand
Madagascar Bulbul (Hypsipetes madagascariensis) Madagascar
Spectacled Bulbul (Pycnonotus erythropthalmos) Thailand
Scaly-breasted Bulbul (Ixodia squamata) Southern Thailand
Grey-bellied Bulbul (Ixodia cyaniventris) Thailand
Black-crested (Rubigula (Pycnonotus melanicterus) flaviventris) Thailand
Flame-throated Bulbul (Rubigula gularis) Tamil Nadu, India
Cream-vented Bulbul (Pycnonotus simplex) Southern Thailand
Red-eyed Bulbul (Pycnonotus brunneus) Southern Thailand
Streak-eared Bulbul (Pycnonotus blanfordi) Thailand
Stripe-throated (Pycnonotus finlaysoni) Thailand
Flavescent Bulbul (Pycnonotus flavescens) Thailand
Yellow-eared Bulbul (Pycnonotus penicillatus) Sri Lanka
Brown-breasted Bulbul (Pycnonotus xanthorrhous) Thailand, Yunnan
Red-whiskered Bulbul (Pycnonotus jocosus) Thailand
Yellow-vented Bulbul (Pycnonotus analis) Thailand
Red-vented Bulbul (Pycnonotus cafer) India
Sooty-headed Bulbul (Pycnonotus aurigaster) Thailand
African Red-eyed Bulbul (Pycnonotus nigricans) Namibia
Common Bulbul (Pycnonotus barbatus) Ethiopia, East Africa
Dark-capped Bulbul (Pycnonotus tricolor) East Africa
Cape Bulbul (Pycnonotus capensis) South Africa
African Yellow Greenbul (Chlorocichla flaviventris) Namibia
Sombre Greenbul (Andropadus importunus) South Africa
Black-headed Bulbul (Pycnonotus atriceps) Thailand
Chinese Bulbul (Pycnonotus sinensis) China
Hairy-backed Bulbul (Tricholestes criniger) Southern Thailand
Ochraceus Bulbul (Alophoixus ochraceus) Thailand
Styan's Bulbul (Pycnonotus taivanus) Taipei China
Puff-throated Bulbul (Alophoixus pallidus) Thailand
Yellow-bellied Bulbul (Alophoixus phaeocephalus) Borneo
Cream-eyed Bulbul (Pycnonotus simplex) Hala Bala National Wildlife Sanctuary Thailand
Collared finchbill (Spizixos semitorques) Anhui, China
Crested Finchbill (Spizixos canifrons) Northern Thailand